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The Songs and Calls of the Great Tit

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The exact timing of breeding varies by a number of factors, most importantly location. Most breeding occurs between January and September; in Europe the breeding season usually begins after March. In Israel there are exceptional records of breeding during the months of October to December. The amount of sunlight and daytime temperatures will also affect breeding timing.

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Great tits are cavity nesters, breeding in a hole that is usually inside a tree, although occasionally in a wall or rock face, and they will readily take to nest boxes.

The nest inside the cavity is built by the female, and is made of plant fibres, grasses, moss, hair, wool and feathers. The number in the clutch is often very large, as many as 18, but five to twelve is more common.

Clutch size is smaller when birds start laying later, and is also lower when the density of competitors is higher. Insularity also affects clutch size, with great tits on offshore islands laying smaller clutches with larger eggs than mainland birds.

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The female undertakes all incubation duties, and is fed by the male during incubation. The timing of hatching, which is best synchronised with peak availability of prey, can be manipulated when environmental conditions change after the laying of the first egg by delaying the beginning of incubation, laying more eggs or pausing during incubation.

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The chicks, like those of all titsare hatched unfeathered and blind. Once feathers begin to erupt, the nestlings are unusual for altricial birds in having plumage coloured with carotenoids similar to their parents in most species it is dun-coloured to avoid predation.

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The nape is yellow and attracts the attention of the parents by its ultraviolet reflectance. This may be to make them easier to find in low light, or be a signal of fitness to win the parents' attention. This patch turns white after the first moult at age two months, and diminishes in size as the bird grows. Feeding of the fledgeling may continue after independence, lasting up to 25 days in chicks from the first brood, but as long as 50 days in the second brood. Inbreeding depression occurs when the offspring produced as a result of a mating between close relatives show reduced fitness.

The reduced fitness is generally considered to be a consequence of the increased expression of deleterious recessive alleles in these offspring. In natural populations of P. The Eurasian sparrowhawk is a predator of great tits, with the young from second broods being at higher risk partly because of the hawk's greater need for food for its own developing young.

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It was originally a specialist tit flea, but the dry, crowded conditions of chicken runs enabled it to flourish with its new host. Although these beetles often remain in deserted nests, they can only breed in the elevated temperatures produced by brooding birds, tits being the preferred hosts.

Incidences of fatal competition are more frequent when nesting times overlap, and climate change has led to greater synchrony of nesting between the two species and flycatcher deaths. Having killed the flycatchers, the great tits may consume their brains.

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The great tit is a popular garden bird due to its acrobatic performances when feeding on nuts or seed. Its willingness to move into nest boxes has made it a valuable study subject in ornithology ; it has been particularly useful as a model for the study of the evolution of various life-history traits, particularly clutch size. The great tit has generally adjusted to human modifications of the environment.

It is more common and has better breeding success in areas with undisturbed forest cover, but it has adapted to human modified habitats. It can be very common in urban areas.

In areas with low frequency background noise pollution, the song has a higher frequency than in quieter areas.

While there have been some localised declines in population in areas with poorer quality habitats, its large range and high numbers mean that the great tit is not considered to be threatened, and it is classed as least concern on the IUCN Red List. From Wikipedia, the free encyclopedia. Linnaeus Parus major. Biology Letters.

Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I.

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Editio decima, reformata in Latin. Holmiae Laurentii Salvii.

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Cassell's Latin Dictionary 5th ed. London: Cassell Ltd. The ornithology of Francis Willughby of Middleton in the county of Warwick, esq. London, United Kingdom: A. Proceedings of the Royal Society B. RA, ed. Check-list of birds of the world. Volume Cambridge, Massachusetts: Museum of Comparative Zoology.

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Biological Journal of the Linnean Society. Archived from the original on 24 July Retrieved 19 February Handbook of the Birds of the World.

Volume Picathartes to Tits and Chickadees. Barcelona: Lynx Edicions. Tits, Nuthatches and Treecreepers.

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Christopher Helm. Zootaxa : The world bird database. Behavioral Ecology and Sociobiology.

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